Interactive Visualizations
Cilia bundle visualizer — explore microscopy simulation of cilia bundles with shape and coordination control.
Brownian particle visualizer — examine different microscopy modalities on fractional Brownian motion.
Filament tracing utility — 'back-of-the-envelope' annotation helper for microscopy movies of cilia-like filaments.
Fish school visualizer — explore collective dynamics of schooling fish each modeled as a dipole swimmer.
Publications (Google Scholar)
F. Ling, Y. Man, and E. Kanso*, Flagellar Wave Reversal via Molecular Motor Asymmetry, in preparation.
F. Ling, A.T. Sahin, B. Miller-Naranjo, S. Aime, D. Roth, N. Tepho, A.S. Vendrame, E. Emken, M. Kiechle, Y. Tesfaigzi, O. Lieleg, and J.C. Nawroth*, High-throughput Mucus Microrheology for Donor and Disease Phenotyping, bioRxiv, 2024.
D. Roth#, A.T. Sahin#, F. Ling, C.N. Senger, E.J. Quiroz, B.A. Calvert, A. van der Does, T.G. Güney, N. Tepho, S. Glasl, A. van Schadewijk, L. von Schledorn, R. Olmer, E. Kanso, J.C. Nawroth*, and A.L. Ryan*, Structure-function Relationships of Mucociliary Clearance in Human Airways, Nature Communications, 2024.
C. Huang, F. Ling, and E. Kanso*, Collective Phase Transitions in Confined Fish Schools, Proceedings of the National Academy of Sciences, 2024.
F. Ling, T. Essock-Burns, M. McFall-Ngai, K. Katija, J.C. Nawroth*, and E. Kanso*, Flow Physics Guides Morphology of Ciliated Organs, Nature Physics, 2024.
H. Hang, Y. Jiao, S. Heydari, F. Ling, J. Merel, and E. Kanso*, Interpretable and Generalizable Strategies for Stably Following Hydrodynamic Trails, bioRxiv, 2023.
Y. Jiao#, F. Ling#, S. Heydari#, N. Heess, J. Merel, and E. Kanso*, Deep Dive into Model-free Reinforcement Learning for Biological and Robotic Systems: Theory and Practice, arXiv preprint, 2023.
A.V. Kanale#, F. Ling#, H. Guo, S.F. Fürthauer, and E. Kanso*, Spontaneous Phase Coordination and Fluid Pumping in Model Ciliary Carpets, Proceedings of the National Academy of Sciences, 2022.
Y. Jiao#, F. Ling#, S. Heydari#, N. Heess, J. Merel, and E. Kanso*, Learning to Swim in Potential Flow, Physical Review Fluids, 2021.
F. Ling and E. Kanso*, Octopus-Inspired Arm Movements, in Bioinspired Sensing, Actuation, and Control in Underwater Soft Robotic Systems, 2020.
Y. Man#, F. Ling#, and E. Kanso*, Cilia Oscillations, Philosophical Transactions of the Royal Society B, 2019.
F. Ling, H. Guo, and E. Kanso*, Instability-driven Oscillations of Elastic Microfilaments, Journal of the Royal Society Interface, 2018.
# equal contribution | * corresponding author
Talk Materials
The 21st Chinese Biophysics Congress (2025) - Quantitative Prediction of Ciliated Organ Function through Unified Fluid Modeling slides
APS talk on Morphological Diversity of Ciliated Organ Flow Physics (2023) slides
Invited talk on Cilia Coordination (2022) - Janelia 4D Cellular Physiology Workshop live recoring, remastered
Lecture on Pattern Formation via Surface Growth (2020) - based on chapters of Alain Goriely slides
Slides introducing 'Active Matter' (2018) - based on Marchetti et al., Rev. Mod. Phys. 2013 slides
DRP: π_3(S^2) and (co)fiber sequences (2015) notes
DRP: What is Persistent Homology? (2015) slides
Random Notes
Hallucinated thoughts on Stein's paradox, human stereotype bias, and AI alignment organized by GPT.
Taking derivatives of eigenvalues gave me more headaches than I thought. Just in case, here's a reminder.
Art-ish
Here is a lifting pun.
Here is a catzilla invasion.
Here is an astronaut confused about gravity.
Here is where I would like to work.
And click here for something completely different:
If I were Springer-Verlag Graduate Texts in Mathematics,
I would be a combination of the following according to the questionnaire...

Robin Hartshorne's Algebraic Geometry

Saunders Mac Lane's Categories for the Working Mathematician

William S. Massey's A Basic Course in Algebraic Topology

W.B.R. Lickorish's An Introduction to Knot Theory

J.-P. Serre's Linear Representations of Finite Groups
So I suppose I should go ahead to keep working on my skills in AG, AT and cats then learn more Knot theory and Rep theory to fulfill my destiny!